Recent literature has split up the variable species H. marmoratus and this one has for the moment become H. swynnertoni. It is not unlikely that further research may still separate this form into a distinct species.
Walking around at 4:30 in the morning has its advantages. This beautiful frog was found calling from an elevated perch on a fern. Relatively common, I saw several of these whenever I passed riparians on the trail. Found during a night hike in Pantiacolla midlevel rainforest, Manu national park, Peru.
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